Lincoln Burrows (played by Dominic Purcell) is falsely convicted of the murder of Terrence Steadman and sentenced to death for the crime. He is surprised to see his younger brother Michael Scofield in the same prison and who is planning to break him out of prison before his execution. He evaded authorities after escaping from prison and has been exonerated of all charges. In the second season of the series, the story continues to follow Lincoln, his brother and other escapees as they try to evade the authorities pursuing them. Away from the prison setting, Michael and Lincoln are frequently featured in scenes together in the first seven episodes. When they fail to retrieve L. J. Burrows before his trial, they decide to head to Utah to find Westmoreland's hidden money. In season 3 he currently free and successfully assisted Michael and James Whistler to break out of Sona. As of season 4, he is now part of Self's covert "A-Team" assembled to bring down The Company.He is also in the very last scene when he visits Michael Scofield's (his brother) grave, and he puts an origami crane on top of the grave so he can remember the time when he made them for him, so he would always know that his brother was caring for him. In season 5, he finds out Michael is still alive and incarcerated in Yemen, and leads an operation to save his brother.
Michael Scofield (played by Wentworth Miller) is the protagonist of the series and younger brother of Lincoln Burrows. He designs an elaborate escape plan and tattoos it to his body to free his older brother who has been framed for murder and sentenced to death via the electric chair. He successfully executes this plan, freeing himself, Lincoln and 6 other convicts, but ultimately fails to evade the police in Panama. In the second season of the series, the story continues to follow Michael, his brother and other escapees as they try to evade the authorities pursuing them. Away from the prison setting, Michael and Lincoln are frequently featured in scenes together in the first seven episodes. When they fail to retrieve L. J. Burrows before his trial, they decide to head to Utah to find Westmoreland's hidden money. Michael and Lincoln travel together, after the fugitives separate to accomplish each of their individual goals. Season 3 is centered around Michael breaking out of prison again, however, this time from the self-governed prison of Sona, situated in Panama. In season 4, he reunites with Sara after discovering her death in season 3 was faked, and she is part of Agent Donald Self's covert "A-Team", which is assembled to bring down The Company. Michael has low latent inhibition, a personality trait which allows him to notice things that others would not. This, along with his genius-level intellect, is what enabled him to break out of both prisons. He also suffers from a tumor of the hypothalamus, which causes nosebleeds, disorientation, and extreme pain. In season 5, he is revealed to be incarcerated in Yemen under a different name; Kaniel Outis. It is gradually revealed that Michael, along with a fellow prisoner named Whip, are undercover CIA operatives, but were betrayed by a mastermind agent known as Poseidon.
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John Abruzzi (played by Peter Stormare; main: season 1; recurring: season 2) is an Italian-American mob boss and the Don of the Abruzzi crime family, who is imprisoned at Fox River with a life sentence. He controls Prison Industry, an internal work program for the convicts, thanks to his connections with Brad Bellick, who is on the payroll of Abruzzi's associates. He appears as an anti-hero where he is less of an antagonist than an unwanted ally to the protagonists. He is convicted based on the testimony of Otto Fibonacci, who saw him order the deaths of two men. Abruzzi plans to exact revenge on Fibonacci, who entered the witness protection program. In the pilot episode, he is approached by Michael Scofield with a PI membership request. Abruzzi at first dismisses the newcomer, but gives him his full attention after he learns that Michael somehow knows where Fibonacci is hiding. Although Michael claims that he will give up the information in exchange for help with his escape plan, Abruzzi has two of Michael's toes cut off trying to extract Fibonacci's location from him. This fails and Abruzzi decides to instead team up with Michael to break out of prison. Although their alliance is initially unstable, with both men being wary of each other, Abruzzi and Michael learn to get along as the season progresses. Abruzzi's status as head of PI proves valuable to Michael's escape plan, as the escape team is able to use work on PI to build an escape tunnel.
Gretchen Louise Morgan (played by Jodi Lyn O'Keefe; seasons 3-4), sometimes operating under the alias Susan B. Anthony, is introduced to the series in the third season as a high-ranking Company field-operative in charge of Whistler's breakout from Sona. Despite claiming to have killed Sara, season 4 revealed she lied and Sara is in fact alive. Gretchen finds herself in The Company's cross hairs after the Whistler/Scylla disaster and later is forced out of necessity to aid Self's "A-Team" against her former employers. She is ruthless and impatient, performing swift executions without hesitation if there is any obstacle to her plan, but is known to have a soft side for the few individuals that she cares about, for example her daughter. She is shown to be able to withstand torture and is exceptionally skilled in martial arts. Her character is mainly an antagonist, and later an occasional, uneasy ally to the protagonists. Gretchen is the main antagonist of the third season. The character is first seen covering scratch wounds on her face with makeup. Later, she meets up with Lincoln at a bar and informs him that The Company has kidnapped both Sara and LJ, and that in order for them to survive, Michael must break James Whistler out of Sona prison. When Lincoln tries to rescue both of them, she claimed to have personally beheaded Sara and warned the brothers not to play games.[6] Gretchen provides assistance to both of the brothers so that their plan would work, such as money for the gravedigger (whom she eventually executed as she suspected that he might report them to the police)[6] and a drug that could help silence a guard.[7] When she finds out that Lincoln is trying to drug the guard much earlier than the hostage exchange time, she becomes furious and moves to behead the fearful LJ,[7] but is stopped when Lincoln reveals the truth. When the escape ultimately fails, Gretchen's supervisor, General Krantz, forced her to carry out the "Bang and Burn" Operation, which involved flying helicopters into Sona to retrieve Whistler. Due to Michael's intervention, this operation fails. Hoping that Lincoln could once again be included into the plan, Gretchen allowed Lincoln to meet his son at an abandoned shack. Michael's confession caused her to be subsequently tortured by waterboarding by Sona guards. However, Gretchen was able to escape from their clutches.[8] James Whistler later revealed that her real name is Gretchen Morgan.[9] Following Michael and Whistler's escape from Sona, Gretchen was able to extract Whistler safely, but she failed to eliminate the brothers. In her final appearance of the season, she is seen driving away in a car with Whistler and Mahone (who had seemingly made a deal with Whistler to work for The Company after the escape).
David "Whip" Martin (played by Augustus Prew; season 5) is introduced as Michael's cellmate in Ogygia in season 5. He is quickly revealed to know that Michael is not Kaniel Outis, and that he and Michael have been breaking people out of prisons around the world for years. Michael repeatedly calls him his (Michael's) whip-hand, hence his nickname. He is loyal to Michael despite not knowing his real name, but as their time in Ogygia goes on and their escape plan doesn't seem to be working, he starts to become more paranoid that Michael will betray him, which only increases when Abu Ramal, the prisoner they are in Ogygia to break out, is released from solitary confinement. Michael repeatedly tells him he will not leave him, but he continues to be frightened, even fighting Michael and forcing a guard to break them up (though this is part of Michael's plan). He confronts Michael during the escape, where Michael finally reveals his plan to leave Ramal behind, satisfying Whip. However, the escape fails, and he, Michael, Ramal and their cellmate Ja are thrown into solitary confinement, where he starts to become paranoid once again as ISIL strikes land closer to the prison. After Michael convinces Ramal to help them escape, he and Ja are released from their cells when Michael gets the keys. They try to persuade Michael to leave Ramal behind, but Michael admits he is necessary. Eventually the four escape the prison along with Sid, another inmate, and formulate a plan to double-cross Ramal, but he turns the tables on them. Whip tells Ramal he was arrested for killing a man in a barfight, and when they are surprised by Lincoln, he takes the opportunity to kill Ramal, which ends up backfiring. With ISIL hunting them, he continues to follow Michael, now knowing his real name, despite two of their plans for escape failing. Eventually the group tries to escape across the desert to Phaecia, a small fishing village, and Whip kills several ISIL members following them. He accompanies Michael and Lincoln to Crete, Greece, and expresses his admiration for an injured Michael. He then accompanies them on to a freighter, arranged by Sucre, back to the States. When they are caught by NAVY Seals, he escapes with Michael, Lincoln and Sucre in a life raft, and is taken by fishing boat to France, from where Michael instead sends him to Chicago instead of New York. During this time, it is revealed that his real name is David Martin, and that he was a survivor in prison, which is why Michael chose him as his partner (and the cover story he told Ramal may have been why he was originally incarcerated). In Chicago, he follows coordinates given by Michael to Lake Michigan, where he meets T-Bag. T-Bag reveals that he is in fact his father, which is the reason why he Michael made him his whip-hand. During the final confrontation with Jacob Ness, aka Poseidon, he attacks him but is shot by A&W, and he dies on his father's arms. As a result, T-Bag kills A&W and goes back to Fox River prison, where he gets revenge for his son when Michael arranges Jacob to be T-Bag cellmate.
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